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The main difference between Skype and VoIP clients is that Skype operates on a peer-to-peer model, rather than the more traditional server-client model. The Skype user directory is entirely decentralized and distributed among the nodes in the network, which means the network can scale very easily to large sizes (currently about 220 million users) without a complex and costly centralized infrastructure.
Skype also routes calls through other Skype peers on the network to ease the traversal of Symmetric NATs and firewalls. This, however, puts an extra burden on those who connect to the Internet without NAT, as their computers and network bandwidth may be used to route the calls of other users.
